:root {
  --ink: #10231f;
  --ink-2: #1f3732;
  --muted: #64746f;
  --paper: #fcfbf7;
  --white: #ffffff;
  --wash: #f2f4ee;
  --line: #d9ded7;
  --green: #14786f;
  --green-dark: #0d5d56;
  --mint: #bce4d9;
  --sand: #eee7d8;
  --radius-sm: 8px;
  --radius-md: 14px;
  --radius-lg: 20px;
  --max-width: 1240px;
}

* { box-sizing: border-box; }
html { scroll-behavior: smooth; scroll-padding-top: 84px; }
body {
  margin: 0;
  color: var(--ink);
  background: var(--paper);
  font-family: Arial, Helvetica, sans-serif;
  font-size: 16px;
  line-height: 1.6;
  -webkit-font-smoothing: auto;
  text-rendering: optimizeLegibility;
}
body, body * {
  -webkit-text-fill-color: currentColor;
  text-shadow: 0 0 0.01px currentColor;
}
button, input, select, textarea { font: inherit; }
button:focus-visible, a:focus-visible, summary:focus-visible {
  outline: 3px solid rgba(20, 120, 111, 0.3);
  outline-offset: 3px;
}
img { max-width: 100%; display: block; }
a { color: inherit; }
sup { font-size: 0.62em; line-height: 0; }

.sr-only {
  position: absolute;
  width: 1px;
  height: 1px;
  padding: 0;
  margin: -1px;
  overflow: hidden;
  clip: rect(0, 0, 0, 0);
  white-space: nowrap;
  border: 0;
}
.skip-link {
  position: fixed;
  top: 12px;
  left: 12px;
  z-index: 100;
  padding: 10px 16px;
  color: white;
  background: var(--ink);
  border-radius: 7px;
  transform: translateY(-140%);
}
.skip-link:focus { transform: translateY(0); }

.section-shell {
  width: min(calc(100% - 56px), var(--max-width));
  margin-inline: auto;
}

.site-header {
  position: sticky;
  top: 0;
  z-index: 50;
  border-bottom: 1px solid rgba(16, 35, 31, 0.12);
  background: rgba(252, 251, 247, 0.92);
  backdrop-filter: blur(18px);
}
.nav-shell {
  width: min(calc(100% - 56px), var(--max-width));
  height: 68px;
  margin-inline: auto;
  display: grid;
  grid-template-columns: 1fr auto 1fr;
  align-items: center;
  gap: 30px;
}
.brand {
  justify-self: start;
  display: flex;
  align-items: center;
  gap: 10px;
  text-decoration: none;
  font-size: 0.88rem;
  font-weight: 800;
  letter-spacing: 0.14em;
}
.brand small {
  padding-left: 10px;
  border-left: 1px solid var(--line);
  color: #75847f;
  font-size: 0.63rem;
  font-weight: 600;
  letter-spacing: 0.1em;
  text-transform: uppercase;
}
.brand-mark {
  width: 28px;
  height: 28px;
  display: flex;
  align-items: center;
  justify-content: center;
  gap: 2px;
  background: var(--ink);
  border-radius: 50%;
}
.brand-mark i { width: 2px; display: block; background: var(--mint); border-radius: 8px; }
.brand-mark i:nth-child(1), .brand-mark i:nth-child(5) { height: 7px; }
.brand-mark i:nth-child(2), .brand-mark i:nth-child(4) { height: 13px; }
.brand-mark i:nth-child(3) { height: 19px; }
.desktop-nav { display: flex; align-items: center; gap: 25px; }
.desktop-nav a {
  position: relative;
  color: #52645f;
  text-decoration: none;
  font-size: 0.79rem;
  font-weight: 700;
}
.desktop-nav a::after {
  content: "";
  position: absolute;
  left: 0;
  right: 0;
  bottom: -8px;
  height: 2px;
  background: var(--green);
  transform: scaleX(0);
  transform-origin: left;
  transition: transform 180ms ease;
}
.desktop-nav a:hover::after, .desktop-nav a[aria-current="true"]::after { transform: scaleX(1); }
.resource-nav { justify-self: end; display: flex; align-items: center; gap: 12px; }
.resource-pending, .resource-link {
  color: #6b7a76;
  font-size: 0.68rem;
  font-weight: 700;
  letter-spacing: 0.08em;
  text-transform: uppercase;
}
.resource-pending { cursor: default; }
.resource-link { color: #3c645b; text-decoration: none; }
.resource-link:hover { color: var(--green-dark); text-decoration: underline; }
.resource-nav > * + * { padding-left: 12px; border-left: 1px solid var(--line); }
.menu-button, .mobile-nav { display: none; }

.hero {
  position: relative;
  padding-block: 70px 0;
}
.hero-meta-line {
  display: flex;
  align-items: center;
  justify-content: space-between;
  gap: 24px;
  padding-bottom: 26px;
  border-bottom: 1px solid var(--line);
}
.eyebrow, .section-index, .card-kicker {
  margin: 0;
  color: var(--green);
  font-size: 0.7rem;
  font-weight: 800;
  letter-spacing: 0.14em;
  text-transform: uppercase;
}
.eyebrow { display: flex; align-items: center; gap: 9px; }
.eyebrow span { width: 22px; height: 2px; background: var(--green); }
.hero-meta-line > p:last-child {
  margin: 0;
  color: #7a8884;
  font-size: 0.66rem;
  font-weight: 700;
  letter-spacing: 0.12em;
  text-transform: uppercase;
}
.hero-copy { padding-block: 42px 40px; }
.hero h1 {
  max-width: 1180px;
  margin: 0;
  font-family: Georgia, "Times New Roman", serif;
  font-size: clamp(3.3rem, 6.35vw, 5.9rem);
  font-weight: 600;
  letter-spacing: -0.05em;
  line-height: 0.98;
}
.hero h1 span { display: block; }
.hero h1 .hero-title-tail {
  margin-top: 20px;
  color: var(--green);
  font-family: inherit;
  font-size: clamp(1.45rem, 2.5vw, 2.15rem);
  font-weight: inherit;
  font-style: normal;
  letter-spacing: -0.025em;
  line-height: 1.2;
}
.hero-deck {
  margin-top: 40px;
  display: grid;
  grid-template-columns: minmax(0, 1.25fr) minmax(280px, 0.75fr);
  gap: clamp(44px, 8vw, 110px);
  align-items: start;
}
.hero-summary {
  max-width: 760px;
  margin: 0;
  color: #344b45;
  font-size: clamp(1.08rem, 1.55vw, 1.28rem);
  line-height: 1.55;
}
.hero-contributions {
  margin: 0;
  padding: 0;
  list-style: none;
  border-top: 1px solid var(--line);
}
.hero-contributions li {
  min-height: 38px;
  display: grid;
  grid-template-columns: 38px 1fr;
  align-items: center;
  border-bottom: 1px solid var(--line);
  color: #394f49;
  font-size: 0.75rem;
  font-weight: 700;
}
.hero-contributions span {
  color: var(--green);
  font-family: ui-monospace, SFMono-Regular, Menlo, monospace;
  font-size: 0.61rem;
}
.hero-footer {
  margin-top: 36px;
  display: grid;
  grid-template-columns: minmax(0, 1fr) auto;
  gap: 48px;
  align-items: end;
}
.authors p { margin: 0; font-size: 0.88rem; font-weight: 700; }
.authors .affiliations {
  display: flex;
  flex-wrap: wrap;
  gap: 2px 15px;
  margin-top: 8px;
  color: #687974;
  font-size: 0.69rem;
  font-weight: 500;
}
.equal-note { margin-top: 4px !important; color: #87938f; font-size: 0.64rem !important; font-weight: 500 !important; }
.hero-actions { display: flex; flex-wrap: wrap; justify-content: flex-end; gap: 8px; }
.button {
  display: inline-flex;
  align-items: center;
  justify-content: center;
  min-height: 42px;
  padding: 0 15px;
  border-radius: 999px;
  font-size: 0.73rem;
  font-weight: 800;
  letter-spacing: 0.025em;
  text-decoration: none;
  cursor: pointer;
}
.button span { margin-left: 7px; font-size: 0.75rem; font-weight: 600; opacity: 0.65; text-transform: uppercase; }
.button:disabled { cursor: default; opacity: 1; }
.button-primary { color: white; background: var(--ink); border: 1px solid var(--ink); }
.button-secondary { color: var(--ink); background: transparent; border: 1px solid #cdd5d0; }
.species-rail {
  display: grid;
  grid-template-columns: repeat(6, 1fr);
  border-top: 1px solid var(--line);
}
.species-rail span {
  position: relative;
  padding: 18px 14px 18px 17px;
  color: #61726d;
  border-right: 1px solid var(--line);
  font-size: 0.68rem;
  font-weight: 700;
}
.species-rail span:last-child { border-right: 0; }
.species-rail span::before {
  content: "";
  position: absolute;
  top: 23px;
  left: 0;
  width: 7px;
  height: 7px;
  background: var(--species);
  border-radius: 50%;
}

.stats-band { color: white; background: var(--ink); }
.stats-grid { display: grid; grid-template-columns: repeat(4, 1fr); }
.stats-grid div { padding: 26px 30px; border-right: 1px solid rgba(255,255,255,.13); }
.stats-grid div:first-child { padding-left: 0; }
.stats-grid div:last-child { border-right: 0; }
.stats-grid strong {
  display: block;
  color: var(--mint);
  font-family: Georgia, "Times New Roman", serif;
  font-size: clamp(1.8rem, 2.7vw, 2.5rem);
  font-weight: 600;
  letter-spacing: -0.035em;
  line-height: 1.05;
}
.stats-grid span {
  color: #a8b6b1;
  font-size: 0.66rem;
  font-weight: 700;
  letter-spacing: 0.08em;
  text-transform: uppercase;
}

.content-section { padding-block: 92px; }
.section-heading {
  display: grid;
  grid-template-columns: minmax(110px, 0.27fr) minmax(0, 1fr);
  gap: 42px;
  align-items: start;
  margin-bottom: 44px;
}
.section-heading h2 {
  max-width: 850px;
  margin: 0;
  font-family: Georgia, "Times New Roman", serif;
  font-size: clamp(2.35rem, 4.4vw, 4.35rem);
  font-weight: 600;
  letter-spacing: -0.045em;
  line-height: 1;
}
.section-heading > div > p {
  max-width: 760px;
  margin: 17px 0 0;
  color: #536761;
  font-size: 1rem;
}
.compact-heading { margin-bottom: 38px; }

.demo-surface {
  overflow: hidden;
  border: 1px solid #cdd5d0;
  border-radius: var(--radius-lg);
  background: white;
  box-shadow: 0 22px 60px rgba(23, 48, 42, 0.07);
}
.species-tabs {
  display: grid;
  grid-template-columns: repeat(6, 1fr);
  padding: 7px;
  gap: 4px;
  background: #e8ede7;
  border-bottom: 1px solid #d4dbd5;
}
.species-tabs button {
  min-height: 44px;
  padding: 6px 9px;
  color: #63746f;
  background: transparent;
  border: 0;
  border-radius: 7px;
  font-size: 0.75rem;
  font-weight: 700;
  cursor: pointer;
}
.species-tabs button:hover { color: var(--ink); background: rgba(255,255,255,.58); }
.species-tabs button[aria-selected="true"] { color: var(--ink); background: white; box-shadow: 0 2px 7px rgba(16, 35, 31, 0.07); }
.anchor-selector { padding: 20px 34px 22px; border-bottom: 1px solid var(--line); background: #f8faf6; }
.anchor-selector-head { display: flex; align-items: baseline; justify-content: space-between; flex-wrap: wrap; gap: 8px 18px; margin-bottom: 12px; }
.anchor-selector-head .context-label { margin: 0; }
.anchor-count { color: #7a8983; font-size: 0.7rem; }
.anchor-buttons { display: flex; flex-wrap: wrap; gap: 8px; }
.anchor-buttons button {
  min-height: 36px;
  padding: 7px 12px;
  color: #5b7168;
  background: white;
  border: 1px solid #d4ded5;
  border-radius: 7px;
  font-size: 0.73rem;
  font-weight: 700;
  line-height: 1.35;
  text-align: left;
  cursor: pointer;
}
.anchor-buttons button:hover { color: var(--green-dark); border-color: #93bcb0; }
.anchor-buttons button[aria-pressed="true"] { color: white; background: var(--green-dark); border-color: var(--green-dark); }
.sample-context {
  display: grid;
  grid-template-columns: 1.35fr 0.65fr;
  gap: 42px;
  padding: 30px 34px;
  background: white;
  border-bottom: 1px solid var(--line);
}
.context-label { margin: 0 0 8px; color: #788782; font-size: 0.75rem; font-weight: 800; letter-spacing: 0.12em; text-transform: uppercase; }
.prompt-placeholder { max-width: 720px; margin: 0; font-family: Georgia, "Times New Roman", serif; font-size: 1.3rem; font-weight: 600; letter-spacing: -0.012em; line-height: 1.4; }
.sample-context dl { margin: 0; display: grid; grid-template-columns: repeat(3, 1fr); gap: 15px; align-self: center; }
.sample-context dl div { padding-left: 12px; border-left: 2px solid var(--line); }
.sample-context dt { color: #81908b; font-size: 0.75rem; font-weight: 700; letter-spacing: 0.09em; text-transform: uppercase; }
.sample-context dd { margin: 3px 0 0; font-size: 0.74rem; font-weight: 700; }
.comparison-tools { display: flex; align-items: center; justify-content: space-between; gap: 12px 20px; padding: 13px 34px; border-bottom: 1px solid var(--line); background: white; }
.comparison-tools[hidden], .sample-spectrogram[hidden] { display: none; }
.comparison-tools p { margin: 0; color: #73857c; font-size: 0.71rem; }
.spectrogram-toggle { flex: none; padding: 5px 0; color: var(--green-dark); background: transparent; border: 0; font-size: 0.72rem; font-weight: 700; cursor: pointer; }
.spectrogram-toggle:hover { text-decoration: underline; }
.audio-comparison { display: grid; grid-template-columns: 1fr 1fr; }
.audio-card { min-width: 0; padding: 30px 34px 32px; background: #fafbf8; }
.audio-card + .audio-card { border-left: 1px solid var(--line); }
.proposed-card { background: #eef8f4; }
.audio-card-head { display: flex; justify-content: space-between; align-items: flex-start; gap: 18px; }
.model-code { color: #74857f; font-size: 0.75rem; font-weight: 800; letter-spacing: 0.13em; }
.audio-card h3 { margin: 5px 0 0; font-size: 1rem; font-weight: 700; }
.status-chip { flex: none; padding: 4px 8px; color: #6e7e79; background: white; border: 1px solid var(--line); border-radius: 999px; font-size: 0.75rem; font-weight: 700; }
.sample-player { display: block; width: 100%; max-width: 500px; height: 48px; margin-top: 24px; } .player-unavailable { display: flex; align-items: center; min-height: 48px; margin-top: 24px; color: #72837d; font-size: 0.82rem; }
.audio-duration { margin: 7px 0 0; color: #7b8d84; font-size: 0.66rem; font-variant-numeric: tabular-nums; }
.audio-duration:empty { display: none; }
.sample-spectrogram { margin: 20px 0 0; }
.sample-spectrogram a { display: block; overflow: hidden; border: 1px solid #dfe5de; border-radius: 8px; background: white; }
.sample-spectrogram img { width: 100%; height: auto; background: white; }
.sample-spectrogram figcaption { margin-top: 7px; color: #74887d; font-size: 0.64rem; }
.player-placeholder {
  margin-top: 25px;
  display: grid;
  grid-template-columns: 40px 1fr 40px;
  align-items: center;
  gap: 13px;
}
.player-placeholder button {
  width: 40px;
  height: 40px;
  padding: 0 0 0 2px;
  color: #91a09b;
  background: white;
  border: 1px solid #ccd5d0;
  border-radius: 50%;
}
.wave-placeholder {
  height: 34px;
  opacity: 0.58;
  background: repeating-linear-gradient(90deg, #9eada8 0 2px, transparent 2px 7px);
  clip-path: polygon(0 48%,3% 37%,6% 65%,9% 42%,12% 18%,15% 79%,18% 37%,21% 62%,24% 44%,27% 24%,30% 72%,33% 36%,36% 58%,39% 29%,42% 70%,45% 45%,48% 32%,51% 61%,54% 21%,57% 78%,60% 41%,63% 66%,66% 45%,69% 28%,72% 63%,75% 38%,78% 57%,81% 31%,84% 69%,87% 46%,90% 34%,93% 60%,96% 39%,100% 50%,100% 53%,96% 61%,93% 40%,90% 66%,87% 54%,84% 31%,81% 69%,78% 43%,75% 62%,72% 37%,69% 72%,66% 55%,63% 34%,60% 59%,57% 22%,54% 79%,51% 39%,48% 68%,45% 55%,42% 30%,39% 71%,36% 42%,33% 64%,30% 28%,27% 76%,24% 56%,21% 38%,18% 63%,15% 21%,12% 82%,9% 58%,6% 35%,3% 63%,0 52%);
}
.proposed-card .wave-placeholder { background: repeating-linear-gradient(90deg, var(--green) 0 2px, transparent 2px 7px); }
.player-placeholder time { color: #7b8b86; font-size: 0.67rem; font-family: ui-monospace, SFMono-Regular, Menlo, monospace; text-align: right; }
.interpretation-note {
  margin-top: 16px;
  display: grid;
  grid-template-columns: 160px 1fr;
  gap: 24px;
  padding: 18px 22px;
  color: #526760;
  background: #eef2ed;
  border-left: 3px solid var(--green);
}
.interpretation-note span { color: var(--ink); font-size: 0.68rem; font-weight: 800; letter-spacing: 0.1em; text-transform: uppercase; }
.interpretation-note p { margin: 0; font-size: 0.82rem; }

.overview-section { border-top: 1px solid var(--line); background: var(--white); }
.abstract-layout { display: grid; grid-template-columns: 1fr 1fr; gap: 78px; }
.abstract-copy p { color: #536761; font-size: 0.95rem; }
.abstract-copy .lead-paragraph { margin-top: 0; color: var(--ink); font-family: Georgia, "Times New Roman", serif; font-size: 1.35rem; font-weight: 600; line-height: 1.45; letter-spacing: -0.015em; }
.contribution-list { border-top: 1px solid var(--line); }
.contribution-list article {
  display: grid;
  grid-template-columns: 38px 160px 1fr;
  gap: 15px;
  padding: 21px 0;
  border-bottom: 1px solid var(--line);
}
.contribution-list span { color: var(--green); font-size: 0.68rem; font-family: ui-monospace, SFMono-Regular, Menlo, monospace; }
.contribution-list h3 { margin: 0; font-size: 0.87rem; }
.contribution-list p { margin: 0; color: #63756f; font-size: 0.78rem; }

#dataset { border-top: 1px solid var(--line); }
.figure-card { margin: 0; border: 1px solid var(--line); border-radius: var(--radius-md); background: white; }
.dataset-figure { padding: 24px; }
.dataset-figure img { width: 100%; }
.figure-card figcaption { margin-top: 15px; color: #71827d; font-size: 0.71rem; }
.figure-card figcaption span { margin-right: 9px; color: var(--ink); font-weight: 800; text-transform: uppercase; }
.species-inventory { display: grid; grid-template-columns: repeat(6, 1fr); margin-top: 22px; border: 1px solid var(--line); border-radius: var(--radius-md); overflow: hidden; background: white; }
.species-inventory article { position: relative; padding: 21px 17px; border-right: 1px solid var(--line); }
.species-inventory article:last-child { border-right: 0; }
.species-inventory article::before { content: ""; position: absolute; top: 0; left: 0; right: 0; height: 3px; background: var(--species); }
.species-inventory span { color: #83918d; font-size: 0.61rem; font-family: ui-monospace, SFMono-Regular, Menlo, monospace; }
.species-inventory h3 { margin: 14px 0 3px; font-size: 0.81rem; line-height: 1.25; }
.species-inventory p { margin: 0; color: #687a74; font-size: 0.69rem; }

.dark-section { color: white; background: var(--ink); }
.inverse-heading .section-index { color: var(--mint); }
.inverse-heading > div > p { color: #aab9b4; }
.method-layout { display: grid; grid-template-columns: minmax(0, 1.3fr) minmax(320px, 0.7fr); gap: 60px; align-items: start; }
.method-figure { margin: 0; padding: 20px; background: white; border-radius: var(--radius-md); }
.method-figure img { width: 100%; }
.method-figure figcaption { margin-top: 12px; color: #6d8079; font-size: 0.69rem; }
.method-figure figcaption span { margin-right: 9px; color: var(--ink); font-weight: 800; text-transform: uppercase; }
.method-steps { margin: 0; padding: 0; list-style: none; border-top: 1px solid rgba(255,255,255,.16); }
.method-steps li { display: grid; grid-template-columns: 44px 1fr; gap: 15px; padding: 23px 0; border-bottom: 1px solid rgba(255,255,255,.16); }
.method-steps span { color: var(--mint); font-size: 0.68rem; font-family: ui-monospace, SFMono-Regular, Menlo, monospace; }
.method-steps h3 { margin: 0 0 6px; font-size: 0.95rem; }
.method-steps p { margin: 0; color: #a2b3ad; font-size: 0.8rem; }
.experimental-setup { margin-top: 40px; padding: 24px; border: 1px solid rgba(255,255,255,.16); border-radius: var(--radius-md); background: rgba(255,255,255,.035); }
.experimental-setup .card-kicker { color: var(--mint); }
.setup-grid { display: grid; grid-template-columns: repeat(4, minmax(0, 1fr)); gap: 18px 24px; margin: 20px 0; }
.setup-grid dt { color: #9db5ab; font-size: 0.63rem; font-weight: 700; letter-spacing: 0.07em; text-transform: uppercase; }
.setup-grid dd { margin: 6px 0 0; color: #e0ece7; font-size: 0.8rem; font-weight: 700; }
.experimental-setup > p:last-child { max-width: 1040px; margin: 0; color: #a5bbb0; font-size: 0.78rem; }

.metric-grid { display: grid; grid-template-columns: repeat(4, 1fr); gap: 10px; margin-bottom: 22px; }
.metric-grid article { padding: 21px; background: #eef2ed; border: 1px solid #dfe5de; border-radius: var(--radius-sm); }
.metric-grid p { display: flex; justify-content: space-between; gap: 10px; margin: 0; color: #65766f; font-size: 0.75rem; font-weight: 700; text-transform: uppercase; letter-spacing: 0.055em; }
.metric-grid p span { color: var(--green); }
.metric-grid strong { display: block; margin-top: 17px; font-family: Georgia, "Times New Roman", serif; font-size: 2rem; font-weight: 600; letter-spacing: -0.035em; line-height: 1; }
.metric-grid small { display: block; margin-top: 7px; color: #7b8b85; font-size: 0.64rem; }
.results-grid { display: grid; grid-template-columns: minmax(0, 1.35fr) minmax(290px, 0.65fr); gap: 22px; }
.table-card, .conditioning-card { min-width: 0; border: 1px solid var(--line); border-radius: var(--radius-md); overflow: hidden; }
.card-title { min-height: 56px; padding: 0 20px; display: flex; align-items: center; justify-content: space-between; gap: 18px; background: #f4f6f2; border-bottom: 1px solid var(--line); font-size: 0.74rem; font-weight: 800; }
.card-title small { color: #7d8d87; font-size: 0.61rem; font-weight: 500; }
.table-scroll { overflow-x: auto; }
table { width: 100%; border-collapse: collapse; font-variant-numeric: tabular-nums; background: white; }
th, td { padding: 16px 18px; border-bottom: 1px solid var(--line); text-align: right; font-size: 0.7rem; white-space: nowrap; }
thead th { color: #71827c; font-size: 0.59rem; text-transform: uppercase; letter-spacing: 0.045em; }
th:first-child { text-align: left; }
tbody th { font-weight: 700; }
tbody tr:last-child th, tbody tr:last-child td { border-bottom: 0; }
.highlight-row { background: #eaf7f2; }
.highlight-row th { color: var(--green-dark); }
.reference-row { color: #81908a; background: #fafbf8; }
.results-table { min-width: 840px; }
.results-table th, .results-table td { padding: 14px; }
.table-note { margin: 0; padding: 14px 20px; color: #6e8278; background: white; border-top: 1px solid var(--line); font-size: 0.68rem; }
.detail-table { margin-top: 22px; overflow: hidden; border: 1px solid var(--line); border-radius: var(--radius-md); background: white; }
.detail-table summary { display: flex; flex-wrap: wrap; align-items: center; gap: 8px 12px; min-height: 57px; padding: 15px 20px; color: var(--ink-2); background: #f4f6f2; font-size: 0.75rem; font-weight: 800; cursor: pointer; list-style: none; }
.detail-table summary::-webkit-details-marker { display: none; }
.detail-table summary::before { content: "+"; width: 12px; color: var(--green); font-size: 1rem; font-weight: 600; }
.detail-table[open] summary { border-bottom: 1px solid var(--line); }
.detail-table[open] summary::before { content: "−"; }
.detail-table summary small { margin-left: auto; color: #7b8f84; font-size: 0.66rem; font-weight: 500; }
.detail-table table { min-width: 620px; }
.conditioning-card { padding: 25px; background: var(--ink); color: white; }
.conditioning-card .card-kicker { color: var(--mint); }
.conditioning-card h3 { margin: 16px 0 24px; font-family: Georgia, "Times New Roman", serif; font-size: 1.55rem; font-weight: 600; line-height: 1.2; letter-spacing: -0.02em; }
.conditioning-card > p:last-child { color: #a8b8b2; font-size: 0.78rem; }
.mini-comparison { display: grid; grid-template-columns: 1fr 1fr; gap: 9px; margin-bottom: 22px; }
.mini-comparison div { padding: 13px; background: rgba(255,255,255,.07); border: 1px solid rgba(255,255,255,.12); border-radius: 7px; }
.mini-comparison span, .mini-comparison small { display: block; color: #98aaa4; font-size: 0.59rem; }
.mini-comparison strong { display: block; margin: 8px 0 3px; font-size: 1.4rem; }
.embedding-figure { margin-top: 22px; display: grid; grid-template-columns: 0.34fr 0.66fr; padding: 30px; align-items: center; gap: 26px; }
.embedding-figure .figure-copy { padding-left: 8px; }
.embedding-figure h3 { margin: 15px 0 12px; font-family: Georgia, "Times New Roman", serif; font-size: 1.4rem; font-weight: 600; line-height: 1.25; letter-spacing: -0.02em; }
.embedding-figure .figure-copy > p:last-child { color: #5f746d; font-size: 0.8rem; }
.embedding-figure img { max-height: 570px; width: 100%; object-fit: contain; }
.embedding-figure figcaption { grid-column: 2; margin: 0; }

.limitations-section { border-top: 1px solid var(--line); }
.scope-grid { display: grid; grid-template-columns: repeat(3, 1fr); border-top: 1px solid var(--line); }
.scope-grid article { padding: 27px 32px 8px 0; border-right: 1px solid var(--line); }
.scope-grid article + article { padding-left: 32px; }
.scope-grid article:last-child { border-right: 0; }
.scope-grid span { color: var(--green); font-size: 0.68rem; font-family: ui-monospace, SFMono-Regular, Menlo, monospace; }
.scope-grid h3 { margin: 19px 0 8px; font-size: 0.95rem; }
.scope-grid p { margin: 0; color: #60746d; font-size: 0.81rem; }

.citation-section { color: white; background: #17342d; }
.citation-layout { display: grid; grid-template-columns: 0.7fr 1.3fr; gap: 80px; align-items: start; }
.citation-layout .section-index { color: var(--mint); }
.citation-layout h2 { margin: 14px 0 10px; font-family: Georgia, "Times New Roman", serif; font-size: clamp(2.2rem, 3.8vw, 3.8rem); font-weight: 600; letter-spacing: -0.04em; line-height: 1.05; }
.citation-layout > div:first-child > p:last-child { color: #a6b7b1; font-size: 0.81rem; }
.citation-card { position: relative; overflow: hidden; background: #0d251f; border: 1px solid rgba(255,255,255,.14); border-radius: var(--radius-md); }
.citation-card pre { margin: 0; padding: 55px 25px 25px; overflow-x: auto; color: #cde8de; font: 0.71rem/1.7 ui-monospace, SFMono-Regular, Menlo, Consolas, monospace; }
.copy-button { position: absolute; top: 12px; right: 12px; padding: 6px 9px; color: #c7d8d2; background: rgba(255,255,255,.06); border: 1px solid rgba(255,255,255,.14); border-radius: 6px; font-size: 0.75rem; font-weight: 700; cursor: pointer; }
.copy-button:hover { background: rgba(255,255,255,.12); }
.copy-status { position: absolute; top: 15px; left: 25px; margin: 0; color: var(--mint); font-size: 0.61rem; }

.site-footer { padding-block: 30px; color: #859a92; background: #0d251f; border-top: 1px solid rgba(255,255,255,.08); }
.footer-grid { display: flex; align-items: flex-end; justify-content: space-between; gap: 24px; }
.footer-grid strong { color: white; letter-spacing: 0.12em; }
.footer-grid p { margin: 3px 0 0; font-size: 0.68rem; }

@media (max-width: 1040px) {
  .nav-shell { grid-template-columns: 1fr auto; }
  .desktop-nav, .resource-nav { display: none; }
  .menu-button {
    justify-self: end;
    width: 40px;
    height: 40px;
    display: grid;
    place-content: center;
    gap: 6px;
    background: transparent;
    border: 1px solid var(--line);
    border-radius: 8px;
  }
  .menu-button span:not(.sr-only) { width: 18px; height: 2px; display: block; background: var(--ink); }
  .mobile-nav { padding: 10px 24px 18px; border-top: 1px solid var(--line); background: var(--paper); }
  .mobile-nav:not([hidden]) { display: grid; grid-template-columns: repeat(5, 1fr); gap: 8px; }
  .mobile-nav a { padding: 9px 5px; text-align: center; text-decoration: none; font-size: 0.75rem; font-weight: 700; }
  .hero-deck { grid-template-columns: 1fr 0.7fr; gap: 42px; }
  .hero-footer { grid-template-columns: 1fr; }
  .hero-actions { justify-content: flex-start; }
  .sample-context { grid-template-columns: 1fr; gap: 26px; }
  .abstract-layout { gap: 46px; }
  .contribution-list article { grid-template-columns: 34px 1fr; }
  .contribution-list p { grid-column: 2; }
  .species-inventory { grid-template-columns: repeat(3, 1fr); }
  .species-inventory article:nth-child(3) { border-right: 0; }
  .species-inventory article:nth-child(-n+3) { border-bottom: 1px solid var(--line); }
  .method-layout { grid-template-columns: 1fr; }
  .setup-grid { grid-template-columns: repeat(2, minmax(0, 1fr)); }
  .metric-grid { grid-template-columns: repeat(2, 1fr); }
  .results-grid { grid-template-columns: 1fr; }
}

@media (max-width: 760px) {
  .section-shell, .nav-shell { width: min(calc(100% - 30px), var(--max-width)); }
  .nav-shell { height: 64px; }
  html { scroll-padding-top: 74px; }
  .brand small { display: none; }
  .mobile-nav:not([hidden]) { grid-template-columns: repeat(2, 1fr); }
  .hero { padding-top: 45px; }
  .hero-meta-line { padding-bottom: 18px; }
  .hero-meta-line > p:last-child { display: none; }
  .hero-copy { padding-block: 30px 32px; }
  .hero h1 { font-size: clamp(3rem, 14vw, 4.4rem); line-height: 0.95; }
  .hero h1 .hero-title-tail { margin-top: 17px; font-size: 1.45rem; line-height: 1.2; }
  .hero-deck { grid-template-columns: 1fr; gap: 28px; margin-top: 28px; }
  .hero-summary { font-size: 1rem; }
  .hero-contributions { max-width: none; }
  .hero-footer { margin-top: 28px; gap: 28px; }
  .authors .affiliations { display: block; }
  .authors .affiliations span { display: block; }
  .hero-actions { display: grid; grid-template-columns: repeat(3, 1fr); }
  .button { padding-inline: 10px; }
  .species-rail { grid-template-columns: repeat(3, 1fr); }
  .species-rail span { border-bottom: 1px solid var(--line); }
  .species-rail span:nth-child(3) { border-right: 0; }
  .species-rail span:nth-child(n+4) { border-bottom: 0; }
  .stats-grid { grid-template-columns: repeat(2, 1fr); }
  .stats-grid div { padding: 21px 10px; border-bottom: 1px solid rgba(255,255,255,.13); }
  .stats-grid div:nth-child(2) { border-right: 0; }
  .stats-grid div:nth-child(3), .stats-grid div:nth-child(4) { border-bottom: 0; }
  .stats-grid div:first-child { padding-left: 10px; }
  .stats-grid strong { font-size: 1.65rem; }
  .stats-grid span { font-size: 0.59rem; }
  .content-section { padding-block: 70px; }
  .section-heading { grid-template-columns: 1fr; gap: 12px; margin-bottom: 32px; }
  .section-heading h2 { font-size: 2.55rem; }
  .section-heading > div > p { font-size: 0.92rem; }
  .species-tabs { display: flex; overflow-x: auto; scroll-snap-type: x mandatory; }
  .species-tabs button { flex: 0 0 auto; min-width: 105px; scroll-snap-align: start; }
  .anchor-selector { padding: 18px 20px; }
  .anchor-selector-head { gap: 5px; }
  .anchor-buttons { gap: 6px; }
  .anchor-buttons button { min-height: 38px; padding-inline: 10px; }
  .sample-context { padding: 25px 20px; }
  .prompt-placeholder { font-size: 1.12rem; }
  .sample-context dl { grid-template-columns: repeat(3, 1fr); gap: 9px; }
  .sample-context dd { font-size: 0.7rem; }
  .comparison-tools { align-items: flex-start; padding: 12px 20px; }
  .comparison-tools p { max-width: 210px; }
  .audio-comparison { grid-template-columns: 1fr; }
  .audio-card { padding: 25px 20px; }
  .audio-card + .audio-card { border-left: 0; border-top: 1px solid var(--line); }
  .audio-card-head { display: block; }
  .status-chip { display: inline-block; margin-top: 11px; }
  .interpretation-note { grid-template-columns: 1fr; gap: 7px; }
  .abstract-layout { grid-template-columns: 1fr; gap: 36px; }
  .abstract-copy .lead-paragraph { font-size: 1.17rem; }
  .dataset-figure { padding: 11px; overflow-x: auto; }
  .dataset-figure img { min-width: 820px; }
  .species-inventory { grid-template-columns: repeat(2, 1fr); }
  .species-inventory article:nth-child(3) { border-right: 1px solid var(--line); }
  .species-inventory article:nth-child(even) { border-right: 0; }
  .species-inventory article:nth-child(4) { border-bottom: 1px solid var(--line); }
  .method-layout { gap: 32px; }
  .experimental-setup { margin-top: 30px; padding: 20px; }
  .setup-grid { gap: 16px; }
  .method-figure { padding: 11px; overflow-x: auto; }
  .method-figure img { min-width: 720px; }
  .metric-grid { grid-template-columns: 1fr 1fr; gap: 8px; }
  .metric-grid article { padding: 17px; }
  .metric-grid strong { font-size: 1.55rem; }
  .embedding-figure { grid-template-columns: 1fr; padding: 20px; }
  .embedding-figure .figure-copy { padding: 0; }
  .embedding-figure figcaption { grid-column: 1; }
  .detail-table summary small { flex-basis: 100%; margin-left: 24px; }
  .scope-grid { grid-template-columns: 1fr; }
  .scope-grid article, .scope-grid article + article { padding: 22px 0; border-right: 0; border-bottom: 1px solid var(--line); }
  .scope-grid article:last-child { border-bottom: 0; }
  .citation-layout { grid-template-columns: 1fr; gap: 30px; }
  .citation-card pre { font-size: 0.64rem; }
  .footer-grid { align-items: flex-start; flex-direction: column; }
}

@media (max-width: 430px) {
  .hero h1 { font-size: clamp(2.65rem, 13vw, 3.35rem); }
  .hero-actions, .metric-grid { grid-template-columns: 1fr; }
  .player-placeholder { grid-template-columns: 38px 1fr; }
  .player-placeholder time { display: none; }
  .species-inventory { grid-template-columns: 1fr; }
  .species-inventory article { border-right: 0 !important; border-bottom: 1px solid var(--line) !important; }
  .species-inventory article:last-child { border-bottom: 0 !important; }
  .mini-comparison { grid-template-columns: 1fr; }
}

@media (prefers-reduced-motion: reduce) {
  html { scroll-behavior: auto; }
  *, *::before, *::after { transition-duration: 0.01ms !important; animation-duration: 0.01ms !important; animation-iteration-count: 1 !important; }
}